The fish crawls onto land. The fin becomes a limb. The ventral sheet of muscle, once a vague slab, now faces a new problem: gravity. The sprawling reptile, say a Hylonomus , needs to stop its leg from splaying out like a wet rag every time it takes a step. Deep in its thigh, the ventral sheet begins to specialize. A thick, round belly of muscle attaches from the pubis (the front of the pelvis) to the femur. It is the puboischiofemoralis internus . Its job: adduction. Pull the leg inward, toward the midline. It is a crude rope, but it works.
Fast-forward 100 million years. The cord has a spine. Fins have sprouted from the flanks of a fish called Eusthenopteron . But the fin is a simple flap, moved by thick blocks of muscle layered on top of each other: dorsal (top) and ventral (bottom). Deep within the ventral wall, a sheet of fibers runs obliquely, helping to pull the fin close to the body. This is not yet the adductor longus, but it is its phantom—a primitive retractor, a keeper of balance in the surge of Devonian tides. origin of adductor longus muscle
Then, a miracle: bipedalism.